ἐν with ὠσίν, through association with such phrases as Aesch. Pers. 605 “βοᾷ δ᾽ ἐν ὠσὶ κέλαδος.” ἐπί with “τῇ ψυχῇ” denotes the seat, and, equally with “ἐν”, here=our ‘in’: cp. Il. 1.55 “τῷ γὰρ ἐπὶ φρεσὶ θῆκε θεά”.
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