Herbert Weir Smyth, A Greek Grammar for Colleges

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[*] 995. Very common are introductory relative clauses forming a nominative predicate of the sentence that follows: ““ὃ δὲ πάντων δεινότατον” but what is most terrible of all” L. 30.29. ἐστί is regularly omitted (944). Such relative clauses are followed by an independent sentence, a clause with ὅτι, by ὅτε γάρ, ὅταν, ὅταν γάρ, <*>. Similarly ““τὸ δ᾽ ἔσχατον πάντων, ὅτι” but what is worst of all” P. Ph. 66d, etc.