Allen and Greenough's New Latin Grammar for Schools and Colleges, chapter 77

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[*] 77. The regular Nominative plural of i-stems is -ēs,¹ but -īs is occasionally found. The regular Accusative plural -īs is common, but not exclusively used in any word. An old form for both cases is -eis (diphthong).

1 The Indo-European ending of the nominative plural, -ĕs (preserved in Greek in consonant stems, as ὄρτυξ, ὄρτυγ-ες), contracts with a stem-vowel and gives -ēs in the Latin i-declension (cf, the Greek plural ὄεις). This -ēs was extended to consonant stems in Latin.

Ginn and Company, 1903.

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