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[*] 90. Most nouns of the Fourth Declension in -us are Masculine. Exceptions: The following are Feminine: acus , anus, colus , domus , īdūs (plural), manus , nurus, porticus , quīnquātrūs (plural), socrus , tribus , with a few names of plants and trees. Also, rarely, penus , specus .
Ginn and Company, 1903.
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